(* = graduate student coauthor, ** = undergraduate student coauthor)


Fisher, J.R.*, Fisher, D.M., Skvarla, M.J., Nelson, W.A.*, Dowling, A.P.G. In review. Revision of torrent mites (Parasitengona: Torrenticolidae: Torrenticola) of the United States and Canada: 90 descriptions, molecular phylogenetics, and a key to species. Zookeys.

Hamilton, F.B.*, Wiedenmann, R.N., Skvarla, M.J.*, Sathyamurthy, R., Fisher, D.M., Fisher, J.R.*, and Dowling, A.P.G. In press. Litter-dwelling ground beetles (Coleoptera: Carabidae) and ground spiders (Araneae: Gnaphosidae) of the Ozark Highlands, USA.

Skvarla, M.J.*, Dowling, A.P.G. 2017. Collecting beetles: Analysis of a single-site data set and comparison of trapping techniques (Coleoptera: Curculionoidea excluding Scolytinae, Carabidae, Buprestidae, Cerambycidae). Journal of Insect Science 17(1); 7: 1-28 doi: 10.1093/jisesa/iew098


Skvarla, M.J.*, Barnes, J.K., Fisher, D.M., Dowling, A.P.G. 2016. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. IV. Asilidae and other Diptera. Biodiversity Data Journal 4: e9977. doi: 10.3897/BDJ.4.e9977

Skvarla, M.J., Fisher, D.M., Dowling, A.P.G. 2016. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. III. Heteroptera (Insecta: Hemiptera). Biodiversity Data Journal 4: e7607. doi: 10.3897/BDJ.4.e7607

Skvarla, M.J., Smith, D.R., Fisher, D.M., Dowling, A.P.G. 2016. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. II. Sawflies (Insecta: Hymenoptera: "Symphyta"). Biodiversity Data Journal 4: e8830. doi: 10.3897/BDJ.4.e8830

Hamilton, F.B.*, Skvarla, M.J.*, Fisher, D.M., Dowling, A.P.G. 2016. Notes on Pterostichus punctiventris (Chaudoir) (Coleoptera: Carabidae) in Arkansas, with a new state record. Coleopterists Bulletin 70(2): 309-313

Cheri, C.**, Fisher, J.R.*, Dowling, A.P.G. 2016. The masked torrent mite, Torrenticola vultumlarvatum sp. n. (Parasitengona: Torrenticolidae): a water mite endemic to the Ouachita Mountains in eastern North America. Acarologia 56(2): 245-256.

Guerra, A.S., Eckerlin, R.P., Dowling, A.P.G., Durden, L.A., Robbins, R., Dittmar, K., Helgen, K.M., Lone, K., Young, H.S. in press. Host-parasite associations in small mammal communities in semiarid savanna ecosystems of East Africa. Journal of Medical Entomology 53(4) 851-860.

O’Neill, J.C.*, Fisher, J.R.*, Nelson, W.A.*, Fisher, D.M., Skvarla, M.J.*, Dowling, A.P.G. 2016. Systematics of testudacarine torrent mites (Parasitengona: Torrenticolidae) with descriptions of 13 new species from North America. Zookeys 582: 13-110 doi:10.3897/zookeys.582.7684


Skvarla, M.J.*, Evans, G.E., Dowling, A.P.G. 2015. Clinical report of Otobius megnini (Acari: Parasitiformes: Argasidae) from a domestic cat (Felis silvestris catus) and first record from Northwest Arkansas. Midsouth Entomologist 8: 64-72

Skvarla, M.J.*, Fisher, D.M.*, Schnepp, K.E., Dowling, A.P.G. 2015. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. I. Select beetles (Coleoptera: Buprestidae, Carabidae, Cerambycidae, Curculionoidea excluding Scolytinae). Biodiversity Data Journal 3: e6832. doi: 10.3897/BDJ.3.e6832

Skvarla, M.J.*, Bertone, M.A., Fisher, J.R., Dowling, A.P.G. 2015. New information about the cypress weevil, Eudociminus mannerheimii (Boheman, 1836) (Coleoptera: Curculionidae: Molytinae): redescription, range expansion, new host records, and a report as a possible causative agent of tree mortality. Coleopterists Bulletin 69(4): 751-757

Skvarla, M.J.*, Tripodi, A.*, Szalanski, A., and Dowling, A.P.G. 2015. New records of Orussus minutus Middlekauff, 1983 (Hymenoptera: Orussidae) represent a significant western range expansion. Biodiversity Data Journal 3: e5793. doi: 10.3897/BDJ.3.e5793

Fisher, J.R.*, Fisher, D.M., Nelson, W.A.*, O’Neill, J.C.*, Skvarla, M.J., Radwell, A.J., Ochoa, R., Bauchan, G., and Dowling, A.P.G. 2015. An integrative description of Torrenticola trimaculata sp. nov. (Parasitengona: Torrenticolidae), a three-spotted, riffle-dwelling mite from eastern North America: morphology, phylogenetics, and taxonomic history of the genus. Acarologia. 55(1): 71-116

Dowling, A.P.G. 2015. The Evolution of parasitism and host associations in mites. In: Morand, S., Krasnov, B.R. & Littlewood, D.T.J. (eds). Parasite Diversity and Diversification - Evolutionary Ecology Meets Phylogenetics. Cambridge University Press. pgs. 265-288.


Fisher, J.R.*, Fisher, D.M., Skvarla, M.J.*, and Dowling, A.P.G. 2014. Pre-ballooning in Ummidia Thorell, 1875 (Araneae: Ctenizidae) from the Interior Highlands, USA: second account from the region and review of mygalomorph ballooning. J. Arachnology. 42(3): 318-321

Skvarla, M.J.*, Larson, J.L., Dowling, A.P.G. 2014. Pitfalls and preservatives: A review. J. Entomol. Soc. Ontario. 145: 15-43.

Skvarla, M.J.* and Dowling, A.P.G. 2014. First report of gynandromorphism in Temnothorax curvispinosus (Mayr, 1886) (Hymenoptera: Formicidae). Proc. Entomol. Soc. Washington. 116(3): 349-353.

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2014. Redescription of Paracaropsis travisi (Baker, 1949) (Trombidiformes: Cheyletidae), with range expansion, additional host records, and reevaluation of cheyletid chaetotaxy based on the sejugal furrow. Acarologia 54(3): 335-345.

Skvarla, M.J.*, Hartshorn, J.A.*, Dowling, A.P.G. 2014. Report on a large collection of Merope tuber Newman, 1838 (Mecoptera: Meropeidae) from Arkansas with notes on collection technique, sex ratio, and male clasper size. Psyche 2014, Article ID 530757, 6 pages.

Wiedenmann, R.N., Dowling, A.P.G., and Barnes, J.K. 2014. What is the value of your insect collection? Amer. Entomologist 60(2): 101-104

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2014. A review of Cunaxidae (Acariformes, Trombidiformes): Histories and diagnoses of subfamilies and genera, keys to world species, and some new locality records. Zookeys 418:1-103


Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2013. On some mites (Acari: Prostigmata) from the Interior Highlands: descriptions of the male, immature stages, and female reproductive systems of Pseudocheylus americanus (Ewing, 1909) and some new state records for Arkansas. Zootaxa 3641(4): 401–419


Skvarla, M.J.* and Dowling, A.P.G. 2012. Some new armascirine cunaxids (Acari: Prostigmata: Cunaxidae) from the Eastern United States. Zootaxa 3194: 1-34

Beard, J.J., Ochoa, R., Bauchan, G.R., Welbourn, W.C., Pooley, C. and Dowling, A.P.G. 2012. External mouthpart morphology in the Tenuipalpidae (Tetranychoidea): Raoiella a case study. Exp. Appl. Acarol. 57: 227-255

Dowling, A.P.G., Ochoa, R., Beard, J.J., Welbourn, W.C., and Ueckermann, E.A. 2012. Phylogenetic investigation of the genus Raoiella (Prostigmata: Tenuipalpidae): Diversity, distribution, and world invasions. Exp. Appl. Acarol. 57: 257-269


Radwell, A.J., Dowling, A.P.G., Smith, I.M., and Kaliki**, V. in press. Kongsbergia robisoni, n. sp. (Acari: Hydrachnidiae: Aturidae) from the interior highlands of North America based on morphology and molecular genetic analysis. Int. J. Acarol.

Skvarla, M.J.*, Fisher, J.R.*, and Dowling, A.P.G. 2011. A new species of Neoscirula (Acari: Prostigmata: Cunaxidae) from the Ozark Highlands. Acarologia 51: 283-293

Beaulieu, F., Dowling, A.P.G., Klompen, H., De Moraes, G.J., and Walter, D.E. 2011. Superorder Parasitiformes Reuter, 1909. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 123-128

Heraty, J., Ronquist, F., Carpenter, J., Hawks, D., Schulmeister, S., Dowling, A.P.G., Murray, D., Munro, J., Deans, A., Wheeler, W., Schiff, N., and Sharkey, M. 2011. Evolution of the Hymenopteran Megaradiation. Molecular Phylogenetics and Evolution 60: 73-88

Fisher, J.R.*, Skvarla, M.J.*, Bauchan, G.R., Ochoa, R., and Dowling, A.P.G. 2011. Trachymolgus purpureus sp. nov., an armored snout mite (Acari: Bdellidae) from the Ozark Highlands: morphology, development, and key to Trachymolgus Berlese. ZooKeys 125: 1-34.

Sharanowski, B.J., Dowling, A.P.G., and Sharkey, M.J. in press. Molecular systematic of Braconidae (Hymenoptera: Ichneumonidae): phylogenetic insights from a multi-gene dataset. Systematic Entomology

Sharkey, M.J., Carpenter, J.M., Vilhelmsen, L., Heraty, J., Liljeblad, J., Dowling, A.P.G., Schulmeister, S., Murray, D., Deans, A.R., Ronquist, F., Krogmann, L., and Wheeler, W.C. 2012. Phylogenetic relationships among superfamilies of Hymenoptera. Cladistics 28: 80-112.

Ochoa, R., Beard, J.J., Bauchan, G.R., Kane, E.C., Dowling, A.P.G., and Erbe, E. 2011. Herbivore exploits chink in armour of host. American Entomologist 57: 26-29.


Dowling, A.P.G., Bauchan, G.R., Beard, J.J., and Ochoa, R. 2010. Scanning electron microscopy vouchers and genomic data from an individual specimen: maximizing the utility of delicate and rare specimens. Acarologia 50: 479-485.

Fisher, J.R.* and Dowling, A.P.G. 2010. Modern methods and technology for doing classical taxonomy. Acarologia 50: 395-409.

Dowling, A.P.G. and OConnor, B.M. 2010. Phylogenetic relationships within the suborder Dermanyssina (Acari: Parasitiformes) and a test of dermanyssoid monophyly. Int. J. Acarol. 36: 299-312.

Roy, L., Dowling, A.P.G., Chauve, C.M. and Buronfosse, T. 2010. Diversity of phylogenetic information according to the locus and the taxonomic level: an example from a parasitic mesostigmatid mite genus. Int. J. Mol. Sci. 11: 1704-1734.

Dowling, A.P.G. and OConnor, B.M. 2010. Phylogeny of Dermanyssoidea (Acari: Parasitiformes) suggests multiple origins of parasitism. Acarologia 50: 113-129.


Dowling, A.P.G. 2009. Ixobioides truncatus (Johnston) comb. nov. (Acari: Mesostigmata: Ixodorhynchidae): a synonymy and redescription. Sys. Appl. Acarol. 14: 216-224.

Bitra, K. Tan, A., Dowling, A.P.G., and Palli, S. R. 2009. Functional characterization of PAS and HES family bHLH transcription factors during the metamorphosis of the red flour beetle, Tribolium castaneum. Gene. 448: 74-87.

Roy, L., Dowling, A.P.G., Chauve, C.M. Lesna I., Sabelis M.W. and Buronfosse, T. 2009. Molecular phylogenetic assessment of host range in Dermanyssus species from France and The Netherlands. Exp. Appl. Acarology 48: 115-142.

Roy, L., Dowling, A.P.G., Chauve, C.M., and Buronfosse, T. 2009. Delimiting species boundaries within Dermanyssus Dugès 1834 (Acari: Mesostigmata) using a total evidence approach. Mol. Phylo. Evo.50: 446-470.


McKern, J.A., Szalanski, A.L., Johnson, D.T. and Dowling, A.P.G. 2008. Molecular phylogeny of Sesiidae (Lepidoptera) Inferred from Mitochondrial DNA sequences. J. Agri. Urban. Enotmol. 25: 165-178.

Quicke, D. L. J., Sharkey, M. J., Laurenne, N. M., and Dowling, A.P.G. 2008. A preliminary molecular phylogeny of the Sigalphinae (Hymenoptera: Braconidae), including Pselaphanus Szepligeti, based on 28S rDNA, with descriptions of new Afrotropical and Madagascan Minanga and Malasigalphus species. J. Nat. Hist. 42: 2703-2719.

Roda, A., Dowling, A.P.G., Welbourn, C., Pena, J., Rodrigues, J.C.V., Hoy, M.A., Ochoa, R., Duncan, R.A. and De Chi, W. 2008. Red palm mite situation in the Caribbean and Florida. Proc. Caribbean Food Crops Soc. 44: 80-87.

Dowling, A.P.G., Ochoa, R., and J. J. Beard. 2008. Preliminary results on phylogeographic patterns of the invasive Red Palm Mite, Raoiella indica (Prostigmata: Tenuipalpidae). EURAAC Congress Proceedings, Montpellier, France. p. 147-154.

Doucet, D., Beliveau, C., Dowling, A.P.G., Simard, J., Feng, Q., Krell, P.J., and Cusson, M. 2008. Prophenoloxidases 1 and 2 from the spruce budworm, Choristoneura fumiferana: Molecular cloning and assessment of transcriptional regulation by a polydnavirus. Arc. Insect Biochem. Phys. 67: 188-201.


Dowling, A. P. G., Bochkov, A., and OConnor, B.M. 2007. Revision of the genus Andreacarus (Acari: Laelapidae) with description of seven new species and a new genus for Australian species formerly placed in Andreacarus. J. Med. Entomol.44: 405-421.

Pitz, K. M.*, Dowling, A. P. G., Sharanowski, B. J., Boring, C. A., Seltmann, K., and Sharkey, M. J. 2007. Phylogenetic relationships among the Braconidae (Hymenoptera: Ichneumonoidea): a reassessment of Shi et al. 2005. Mol. Phlyo. Evol.43: 338-343.

Cusson, M., Beliveau, C., Sen, S. E., Vandermoten, S., Rutledge, R. G., Stewart, D., Francis, F., Haubruge, E., Rehse, P., Huggins, D. J., Dowling, A. P. G., and Grant, G. H. 2006. Characterization and tissue-specific expression of two lepidopteran farnesyl diphosphate synthase homologs: implications for the biosynthesis of ethyl-substituted juvenile hormones. TProtein 65: 742-758.

Dowling, A. P. G. 2006. Mesostigmatid mites as parasites of small mammals. In: Morand, S., Krasnov, B.R. & Poulin, R. (eds). Micromammals and macroparasites: From evolutionary ecology to management. Springer-Verlag, Tokyo.

Reeves, W.K., Dowling, A. P. G., and Dasch, G. A. 2006. Rickettsial agents from parasitic Dermanyssoidea (Acari: Mesotigmata). Exp. App. Acarology 38: 181-188.

Brooks, D. R., Dowling, A. P. G., van Veller, M. G. P., and Hoberg, E. P. 2004. Ending a decade of deception: a valiant failure, a not-so-valiant failure, and a success story. Cladistics 20: 32-46.

Klimov, P. B., Lekveishvili, M., Dowling, A. P. G., and OConnor, B. M. 2004. Multivariate analysis of morphological variation in two cryptic species of Sancassania (Acari: Acaridae) from Costa Rica. Ann. Entomol. Soc. Am. 97: 322-345.

Dowling, A. P. G., van Veller, M. G. P., Hoberg, E. P., and Brooks, D. R. 2003. A priori and A posteriori methods in comparative evolutionary studies of host-parasite associations. Cladistics 19: 240-253.

Dowling, A. P. G. 2002. Testing the accuracy of treemap and brooks parsimony analyses of coevolutionary patterns using artificial associations. Cladistics 18:416-435.